Hendricks was better wrestler, physically stronger, better top control. You go back and watch some of his earlier fights, Hendricks was pretty good at striking from the clinch and from punching range was a menace, even outside of punching range great at closing distance with his strikes and takedown entries.
The only thing Usman and Colby really ever had on Hendricks was cardio, I mean I think Usmans got a much nicer jab but over all not really as good of striking. I'd take Hendricks to beat Colby pretty easily in a 3 rounder, in a 5 rounder it could be closer but I suspect Hendricks could secure the first 3, real chance he finishes or 10-8s Colby one of the rounds even. I honestly think Koscheck vs Colby is a more interesting question.