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It's all about the S.A.I.D (specific adaptation to imposed demands) principle. The physiology of a high level runners has adapted to running, not fighting. Take Diaz, Usman, Colby and all of the best gas tanks in MMA and they won't be able to keep up with Mo Farah on a 10 mile run, at all. That doesn't mean they are out of shape, it just means they haven't physiologically adapted to the specific movement of running like Mo Farah has.
ABSOLUTELY!!
muscle groups have fast and slow twitch fibers and the body makes physiological adaptations that are very activity specific
grappling is a VERY different activity than running and is clearly far more demanding however a world class grappler could NEVER compete at the elite levels of distance running just as a world class runner would get smoked on the mat
combat sports have a phenomenal learning curve as well where as running is a very natural activity that doesn't require years of practice to achieve proficiency
whenever a combat athlete is out on the road "running for cardio" they are giving away valuable mat time where they could be learning new skills at the same time as building excellent sport specific cardio
running for cardio will convert your fast to slow twitch fibers, lower your testosterone and has a high chance of injury plus it is an activity 100% devoid of martial arts skills