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I'd say its because Hughes had a bad night his second fight against GSP -- that's common in sports. The better long run side sometimes loses. Hughes was good enough to finish GSP once. That makes him, at least relative to GSP, a better fighter than Shields and Hendricks. Hughes also was a champion for a long time. That makes him a better fighter than Shields and Hendricks by just about every other measure. So beating Hughes was a much bigger achievement for GSP than beating Shields and Hughes -- and that's what's meant by best win.
If you're going by GSP's closest (ie most competitive) fights, then the first fight with Penn was the closest. Its also his second most damaging fight (after Serra) -- after the first Penn fight GSP went straight to the hospital, no press conference. After Hendricks GSP went straight to the press conference -- ie Hendricks didn't do as much damage to GSP as Penn did.
And GSP's fight with Karo was closer than his fight with Shields -- Karo, twice got close to getting a submission, Shields never came anywhere close to that. So I'd say GSP's two most competitive fights were Hendricks (GSP should have lost, but it hangs on one very close round, the first round) and the first BJ Penn fight (and I'd argue that GSP should have lost that one too).
Nothing you described makes Hughes a better fighter than Shields or Hendricks. Hughes, in his prime, beat a23 year old GSP who had 7 fights.
Hughes is an mma fossil and would lose to Shields and get his head taken off by Hendricks.